Thanks for the clarification, that is what I had thought. Here is a link to an interesting article on hydroxymethylation and how it regulates retinal neurogenesis in a cell non-autonomous way by regulating Wnt and Notch pathways in zebrafish. You may have already seen it, but thought it may provide some mechanistic insight to what is going on with the paternal DNA during fertilization. https://www.ncbi.nlm.nih.gov/pubmed/28926578
Cheers!